//---------------------------------------------------------------------

        /// <summary>
        /// Grows an individual cohort for a year, incrementing its age by 1
        /// and updating its biomass for annual growth and mortality.
        /// </summary>
        /// <param name="index">
        /// The index of the cohort to grow; it must be between 0 and Count - 1.
        /// </param>
        /// <param name="site">
        /// The site where the species' cohorts are located.
        /// </param>
        /// <returns>
        /// The index of the next younger cohort.  Note this may be the same
        /// as the index passed in if that cohort dies due to senescence.
        /// </returns>
        public int GrowCohort(int index,
                              ActiveSite site,
                              bool annualTimestep)
        {
            Debug.Assert(0 <= index && index <= cohortData.Count);
            Debug.Assert(site != null);

            Cohort cohort = new Cohort(species, cohortData[index]);

            Debug.Assert(cohort.WoodBiomass <= cohort.WoodBiomass + cohort.LeafBiomass);
            Debug.Assert(cohort.LeafBiomass <= cohort.WoodBiomass + cohort.LeafBiomass);

            //  Check for senescence
            if (cohort.Age >= species.Longevity)
            {
                RemoveCohort(index, cohort, site, null);
                return(index);
            }

            if (annualTimestep)
            {
                cohort.IncrementAge();
            }

            float[] biomassChange = Cohorts.BiomassCalculator.ComputeChange(cohort, site);

            Debug.Assert(-(cohort.WoodBiomass + cohort.LeafBiomass) <= biomassChange[0] + biomassChange[1]);  // Cohort can't loss more biomass than it has


            cohort.ChangeWoodBiomass(biomassChange[0]);
            cohort.ChangeLeafBiomass(biomassChange[1]);

            if (cohort.WoodBiomass + cohort.LeafBiomass > 0)
            {
                cohortData[index] = cohort.Data;
                return(index + 1);
            }
            else
            {
                RemoveCohort(index, cohort, site, null);
                return(index);
            }
        }
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        //---------------------------------------------------------------------

        private void RemoveCohort(int index,
                                  ICohort cohort,
                                  ActiveSite site,
                                  ExtensionType disturbanceType)
        {
            /*if (isDebugEnabled)
             *  log.DebugFormat("  cohort removed: {0}, {1} yrs, {2} Mg/ha ({3})",
             *                  cohort.Species.Name, cohort.Age, cohort.Biomass,
             *                  disturbanceType != null
             *                      ? disturbanceType.Name
             *                      : cohort.Age >= species.Longevity
             *                          ? "senescence"
             *                          : cohort.Biomass == 0
             *                              ? "attrition"
             *                              : "UNKNOWN");
             *
             */
            cohortData.RemoveAt(index);
            Cohort.Died(this, cohort, site, disturbanceType);
        }
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        //---------------------------------------------------------------------

        /// <summary>
        /// Computes how much a disturbance damages the cohorts by reducing
        /// their biomass.
        /// </summary>
        /// <returns>
        /// The total of all the cohorts' biomass reductions.
        /// </returns>
        public int DamageBySpecies(IDisturbance disturbance)
        {
            //  Go backwards through list of cohort data, so the removal of an
            //  item doesn't mess up the loop.
            isMaturePresent = false;
            int totalReduction = 0;

            for (int i = cohortData.Count - 1; i >= 0; i--)
            {
                Cohort  cohort    = new Cohort(species, cohortData[i]);
                float[] reduction = disturbance.ReduceOrKillMarkedCohort(cohort);

                if (reduction[0] + reduction[1] > 0)
                {
                    totalReduction += (int)(reduction[0] + reduction[1]);
                    if (reduction[1] < cohort.LeafBiomass)
                    {
                        cohort.ChangeLeafBiomass(-reduction[1]);
                        cohortData[i] = cohort.Data;
                    }
                    if (reduction[0] < cohort.WoodBiomass)
                    {
                        cohort.ChangeWoodBiomass(-reduction[0]);
                        cohortData[i] = cohort.Data;
                    }
                    else    // Assume that if all wood biomass lost, the cohort is dead
                    {
                        RemoveCohort(i, cohort, disturbance.CurrentSite,
                                     disturbance.Type);
                        cohort = null;
                    }
                }
                if (cohort != null && cohort.Age >= species.Maturity)
                {
                    isMaturePresent = true;
                }
            }
            return(totalReduction);
        }